About Growing Degree Hours | Predicting Harvest Date | Harvest Prediction Model | References
- Growing Degree Hours & GDH30
- Why are GDH30 & Harvest Timing Related?
- Why are GDH30 & Fruit Size Related?
- How can growers use GDH30 figures?
Growing degree hours (GDH) are a weighted measurement of the temperatures to which a tree is exposed, based on the idea that trees have an optimum temperature for growth and that temperatures far above or below that optimum temperature have a greater impact than temperatures just above or just below it. UC Davis’s Harvest Prediction Model calculates GDH values based on the equation of Anderson et al. (1986) and temperature readings collected by the more than 100 CIMIS weather stations throughout the state.
Analysis of the accumulated growing degree hours for the first thirty days after full bloom (GDH30) have shown the amount of accumulation to be related to the number of days between full bloom date, reference date, and harvest date (Marra et al., 2002). Higher GDH30 values correlate with less time between full bloom and reference date in peaches (Lopez and DeJong, 2007) and less time between full bloom and harvest date in peaches, nectarines, plums and prunes (Ben Mimoun and DeJong, 1999; DeBuse et al, In press). Higher GDH30 accumulations with no change in management practices also result in smaller fruit at harvest (Lopez and DeJong, 2007). The Harvest Prediction Model enables growers to better integrate knowledge of this relationship into their management practices.
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Why are GDH30 & Harvest Timing Related?
Peach fruit have been shown to have two phases of development. During phase I, the relative growth rate (rate of unit dry weight increase per unit dry weight per unit time, RGR) of the fruit is decreasing at a logarithmic rate. There is then a shift to a stable, linear RGR which constitutes phase II. During phase I, cells are dividing and differentiating. During phase II, cells are expanding and maturing. The timing of this phase change is genetically regulated but also affected by how warm or cold a spring is, and shortening or lengthening of phase I affect the timing of maturity, and thus the harvest date (DeJong & Goudriaan, 1989).
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Why are GDH30 & Fruit Size Related?
Trees are made up of a number of individual organs – fruits, shoots, leaves, etc. Each organ’s potential for growth and pattern of development is based on the genetics of that organ type. Once the potential for growth of that organ is activated by signals within the tree and/or from the environment, the actual "realized" growth is a result of environmental conditions (temperature, light, etc), resource availability (carbohydrates, nutrients) and competition with other organs for those resources. How competitive an organ is, its "sink strength," is dictated by that organ’s relative distance from the source of carbohydrates and the organ’s efficiency in unloading those resources from the supply stream (DeJong, 1999).
During the first stage of growth in peaches, fruit growth is limited by competition from other organs (DeJong and Grossman, 1995). The respiration rate of both vegetative organs and fruit also increases with increased temperature, approximately doubling when temperatures increase by 10° C (Grossman and DeJong, 1994; Pavel and DeJong, 1993). The demands of growth and respiration under higher temperatures quickly add up - between 20 and 30 days after full bloom, the daily potential rate of fruit respiration and growth, and subsequent demand for carbohydrates, can be 5 to 10 times higher during a warmer spring than during a cooler spring (Lopez and DeJong., 2008). These increased respiratory and growth demands come at a time of low photosynthetic fixation because leaves are still small and days are still short (Grossman and DeJong, 1994).Thus not only do fruit have less time for growth because of an earlier growth phase change and an earlier harvest, they also have to deal with increased competition for assimilates from increased maintenance respiration.
Source demands are highest on trees with moderate and high loads of fruit (DeJong and Grossman 1995). Decreasing competition for resources by thinning is thus an obvious solution, but the timing of that thinning is critical to its success. Limiting resources during phase I, cell division, irreparably decreases fruit size because fewer cell divisions leads to fewer cells, decreasing the maximum relative growth rate a fruit can achieve (Grossman and DeJong, 1995b). While thinning fruit at a variety of points throughout the growing season can boost fruit relative growth rate to near its maximum potential given normal temperatures, the absolute growth rate (rate of unit dry weight increase per unit time) cannot be recovered, resulting in smaller fruit (Grossman and DeJong, 1995b). This situation can be likened to a savings account: even given the same interest rate, an account with a larger initial investment will yield more than that with a smaller investment. And though fruit growth rate does increase with increased GDH accumulation, it does not increase enough to compensate for the shortened growing season, resulting in smaller fruit after warmer springs (Lopez and DeJong, 2007). Thus, though thinning is traditionally done 30 days after bloom for early-maturing cultivars and 60 days for late-maturing varieties, these activities should be undertaken sooner with higher GDH30 accumulation.
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How Can Growers Use GDH30 Figures?
Understanding the affects of increased GDH30 empowers growers to adjust their management practices to at least partially counteract the effects of warmer springs. Anticipating earlier harvests can allow growers to organize harvest logistics. Anticipating smaller fruit under normal management, growers can thin fruit earlier to provide more resources on a per-fruit basis over the growing season. As a rule of thumb, 6,000 GDH30 is a critical turning point for fruit size reduction and earlier harvest dates (Lopez and DeJong, 2007; Lopez and DeJong, 2008). When GDH30 goes above this threshold, growers should thin earlier in the season. More exact prediction models for harvest date have been developed for a number of stone fruit cultivars based on the relationship between GDH30 and harvest date (Ben Mimoun and DeJong, 1999; Day et al., 2008).
For guidelines on predicting harvest date, proceed to the next page Predicting Harvest Date. In subsequent pages, the Harvest Prediction Model will help you find your GDH30, which is necessary for calculations.